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Research: Plant systematics and population genetics, chromosome evolution, phylogenetic and comparative analysis
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Sympatric parallel diversification of major oak clades in the Americas and the origins of Mexican species diversity.
Historical introgression among the American live oaks and the comparative nature of tests for introgression.
The evolution and diversification of the red oaks of the California Floristic Province (Quercus section Lobatae, series Agrifoliae).